The arrival of Pokémon Soul Silver and Pokémon Heart Gold dominated Japan last week.
M-Creata data counted 723,000 and 720,000 sales, respectively, as the pair took first and second place.
Third-place Tamodachi Collection sold a puny 74,000 units to break a lifetime total of 1m, while Dragon Quest IX shifted 37,000 copies to take fourth and soldier on to the 4m milestone.
Pokémon's effect on the region meant PS3 dominance - brought about by the launch of the PS3 Slim - lasted only a week. DSi and DS Lite combined to sell 77,677 units for the week ending 13th September, while Sony's flagship console managed 55,344 sales.
That was still way ahead of PSP and Wii, which posted 18,375 and 17,568 sales apiece - and a mountain for Xbox 360 to climb with 7568 sales. PS2 sat at the bottom of the pile with 2612 sales.
There's little else of note, although budget editions of Call of Duty 4 debuted at 12 and 14 on PS3 and 360.