UPDATE 11.16pm: Actor Todd Stashwick, who provided the voice of an unrevealed character in the Uncharted 4 teaser, has also left the project as his character has been recast.
The actor/writer confirmed the casting changes earlier today to IGN.
Last week it was announced that Stashwick would be joining Uncharted creator Amy Hennig at Visceral Games where they'd reunite to work on an upcoming Star Wars game.
ORIGINAL STORY: 6.58 pm: The Last of Us' lead artist, Nathan Wells, has departed Naughty Dog to join Giant Sparrow, the developer behind The Unfinished Swan.
Wells made the announcement on Twitter, where he noted that he "joined the wild, wacky, and ultimately brilliant folks at #GiantSparrow on April 3rd." This was shortly before Naughty Dog announced The Last of Us Remastered, the spruced up PS4 rendition of its post-apocalyptic adventure.
Prior to working on The Last of Us, Wells worked at Irrational Games for 13 years, where he served as art director on BioShock Infinite.
It's unclear what Wells will be working on at Giant Sparrow, but the indie developer teased its upcoming project, codenamed Edith Finch, in March 2013. It will be the second part of the studio's three-game deal with Sony, and the developer noted that Finch won't be a sequel to Swan, but "it's not a million miles away either."
"If you like surreal experiences and games that aren't much like anything you've ever played before then you'll probably like it," Giant Sparrow said at the time.
Wells marks the third major departure from Naughty Dog recently, as Uncharted creative director Amy Hennig left the studio in early March to join Dead Space developer Visceral Games on its upcoming Star Wars title, while Uncharted 3 & 4 director Justin Richmond left to work at League of Legends developer Riot Games.